21. Hartline, Haldan Keffer hartline, haldan keffer. (b. Dec. 22, 1903, Bloomsburg, Pa., USd. March 17,1983, Fallston, Md.), American physiologist who was a cowinner (with George http://www.britannica.com/nobel/micro/260_49.html

Hartline, Haldan Keffer (b. Dec. 22, 1903, Bloomsburg, Pa., U.S.d. March 17, 1983, Fallston, Md.), American physiologist who was a cowinner (with George Wald and Ragnar Granit ) of the 1967 Nobel Prize in Physiology or Medicine for his work in analyzing the neurophysiological mechanisms of vision. Hartline began his study of retinal electrophysiology as a National Research Council Fellow at Johns Hopkins University, Baltimore, receiving his M.D. in 1927. After attending the universities of Leipzig and Munich as an Eldridge Johnson traveling research scholar, he became professor of biophysics and chairman of the department at Johns Hopkins in 1949. He joined the staff of Rockefeller University, New York City, in 1953 as professor of neurophysiology. Hartline investigated the electrical responses of the retinas of certain arthropods, vertebrates, and mollusks because their visual systems are much simpler than those of humans and are thus easier to study. He concentrated his studies on the eye of the horseshoe crab ( Limulus polyphemus ). Using minute electrodes in his experiments, he obtained the first record of the electrical impulses sent by a single optic nerve

22. Hartline, Haldan Keffer --  Britannica Concise Encyclopedia - The Online Encyc hartline, haldan keffer body US physiologist. Biographies of Ragnar Granit,haldan keffer hartline, and George Wald. Includes text of the presentation http://www.britannica.com/ebc/article?tocId=9366668&query=depressed&ct=

23. Hartline, Haldan Keffer haldan keffer hartline was born in Bloomsburg, Pennsylvania, on December 22nd, 1903.His parents were teachers there in the State Normal School (now http://www.cartage.org.lb/en/themes/Biographies/MainBiographies/H/Hartline/Hartl

Hartline, Haldan Keffer Haldan Keffer Hartline was born in Bloomsburg, Pennsylvania, on December 22nd, 1903. His parents were teachers there in the State Normal School (now Bloomsburg State College) where he received his early education. His father, Daniel S. Hartline, was Professor of Biology, but a man whose wide interests also included Astronomy and Geology. It was through his father that Keffer became interested in Natural Sciences. Keffer Hartline attended Lafayette College in Easton, Pennsylvania, graduating in 1923 (B. Sc.). His college teacher of biology, Beverly W. Kunkel, encouraged him to undertake research; his first scientific paper concerned visual responses of land isopods. Summers at the Marine Biological Laboratory at Woods Hole added to his biological training; there he was especially influenced by Jacques Loeb, Selig Hecht, and Merkel H. Jacobs. In the autumn of 1923 he entered the Johns Hopkins School where he was encouraged to continue his research interest in vision in the Department of Physiology under E. K. Marshall and C. D. Snyder. Dr. Snyder let him use his Einthoven string galvanometer with which Hartline undertook the study of the retinal action potential using frogs, decerebrate cats and rabbits. He learned to obtain electroretinograms from intact animals, and recorded clearly recognizable retinal action potentials from human subjects. He also used intact insects for quantitative studies.

24. Encyclopedia: Haldan Keffer Hartline Other descriptions of haldan keffer hartline. haldan keffer hartline (December22, 1903 March 17, 1983) was an American physiologist who was a cowinner http://www.nationmaster.com/encyclopedia/Haldan-Keffer-Hartline

Supporter Benefits Signup Login Sources ... Pies Related Articles People who viewed "Haldan Keffer Hartline" also viewed: George Wald Horseshoe crab Arthropod Erik Estrada What's new? Our next offering Latest newsletter Student area Lesson plans Recent Updates Zheng He Zeus Yukon general election, 2006 Yoji Biomehanika ... More Recent Articles Top Graphs Richest Most Murderous Most Taxed Most Populous ... More Stats Encyclopedia: Haldan Keffer Hartline Updated 217 days 5 hours 3 minutes ago. Other descriptions of Haldan Keffer Hartline Haldan Keffer Hartline December 22 March 17 ) was an American physiologist who was a cowinner (with George Wald and Ragnar Granit ) of the Nobel Prize in Physiology or Medicine for his work in analyzing the neurophysiological mechanisms of vision. Hartline began his study of retinal electrophysiology as a National Research Council Fellow at Johns Hopkins University Baltimore , receiving his M.D. in . After attending the universities of Leipzig and Munich as an Eldridge Johnson traveling research scholar, he became professor of biophysics and chairman of the department at Johns Hopkins in . He joined the staff of Rockefeller University New York City , in as professor of neurophysiology.

25. Encyclopedia: Retina haldan keffer hartline (December 22, 1903 March 17, 1983) was an Americanphysiologist who was a cowinner (with George Wald and Ragnar Granit) of the 1967 http://www.nationmaster.com/encyclopedia/Retina

Supporter Benefits Signup Login Sources ... Pies Related Articles People who viewed "Retina" also viewed: Rod cell Photoreceptor cell Cone cell Eye ... Ciliary body What's new? Our next offering Latest newsletter Student area Lesson plans Recent Updates Zheng He Zeus Yukon general election, 2006 Yoji Biomehanika ... More Recent Articles Top Graphs Richest Most Murderous Most Taxed Most Populous ... More Stats Encyclopedia: Retina Updated 41 days 14 hours 19 minutes ago. Other descriptions of Retina Human eye cross-sectional view. Courtesy NIH National Eye Institute . Many animals have eyes different from the human eye. The retina is a thin layer of cells at the back of the eyeball of vertebrates and some cephalopods ; it is the part of the eye which converts light into nervous signals. An eye is an organ that detects light. ... The National Institutes of Health is an institution of the United States government which focuses on medical research. ... The National Eye Institute (NEI) is one of the US National Institutes of Health that was established in 1968. ... An eye is an organ that detects light. ...

26. George Wald -- Facts, Info, And Encyclopedia Article (Click link for more info and facts about haldan keffer hartline) haldan kefferhartline haldan keffer hartline Julius Axelrod Antonio Damasio http://www.absoluteastronomy.com/encyclopedia/G/Ge/George_Wald.htm

George Wald [Categories: 1997 deaths, 1906 births, Nobel Prize in Physiology or Medicine winners, American scientists, Neuroscientists] George Wald (November 18, 1906–April 12, 1997) was an (A native or inhabitant of the United States) American (A person with advanced knowledge of one of more sciences) scientist who is best known for his work with pigments in the (The light-sensitive membrane covering the back wall of the eyeball; it is continuous with the optic nerve) retina . He won a share of the 1967 (Click link for more info and facts about Nobel Prize in Physiology or Medicine) Nobel Prize in Physiology or Medicine with (Click link for more info and facts about Haldan Keffer Hartline) Haldan Keffer Hartline and (Click link for more info and facts about Ragnar Granit) Ragnar Granit Research As a postdoctoral researcher, Wald discovered that (Any of several fat-soluble vitamins essential for normal vision; prevents night blindness or inflammation or dryness of the eyes) vitamin A was a component of the retina. His further experiments showed that when the pigment (A red photopigment in the retinal rods of vertebrates; dissociates into retinene by light)

27. EXPLORE Haldan Keffer Hartline - Dictionary Of Famous People haldan keffer hartline. Article from free online dictionary of famous people withover 130000 biographies. http://www.dictionaryofeverything.com/explore/723/Haldan_Keffer_Hartline.html

28. NASA Neurolab Web: Mission Home Page haldan keffer hartline (19031983), American biophysicist, first determined themechanics of how the nerves of the visual system receive information and http://neurolab.jsc.nasa.gov/hartline.htm

Spotlight on Neuroscience Haldan Keffer Hartline Haldan Keffer Hartline (1903-1983), American biophysicist, first determined the mechanics of how the nerves of the visual system receive information and transfer it to the brain. He was born in Bloomsburg, Pennsylvania. Hartline worked with horseshoe crabs and frogs to investigate the mechanisms of vision. His discoveries convinced researchers that the retina (the innermost layer of the eye that is light sensitive) and optic nerves themselves process many nerve signals before the signals are transmitted to the brain. Besides providing new knowledge about the neurophysiological mechanisms of vision, Hartline's research contributed to advances in night-vision, pattern-recognition, and motion-detection devices. He shared the 1967 Nobel Prize in physiology or medicine with Swedish neurophysiologist Ragnar Granit and American biochemist George Wald for their work on the biochemical and electrical processes of vision. Curator: Julie Heath and Jacque Havelka Responsible NASA Official: Katherine Newkirk Updated 9 March 1998

29. International: Italiano: Salute: Medicina: Medici E Ricercatori - Open Site hartline, haldan keffer (0); Hartwell, Leland (0); Hench, Philip Showalter (0);Hershey, Alfred (0); Hess, Walter Rudolf (0) http://open-site.org/International/Italiano/Salute/Medicina/Medici_e_Ricercatori

Open Site The Open Encyclopedia Project Pagina principale Aggiungi Contenuti Diventa Editore In tutta la Directory Solo in Medicina/Medici_e_Ricercatori Top International Italiano Salute ... Medicina : Medici e Ricercatori Ippocrate Dioscoride Galeno Teofrasto ... Zinkernagel, Rolf Vedi anche: International: Italiano: Societ : Strutture Sociali: Persone: Biografie Questa Categoria ha bisogno di un Editore - Richiedila Open Site Code 0.5.3 robot company. Visit our sister sites dmoz.org mozilla.org chefmoz.org musicmoz.org ... modifica

30. Der Nobelpreis Für Physiologie Oder Medizin: Haldan Keffer Hartline Granit und George Wald ). haldan keffer hartline *1903, † 1983. PAGERANK SEO. http://www.nobelpreis.org/medizin/hartline.htm

vor suchen Home Chemie ... Wirtschaft Haldan Keffer Hartline (USA) "Für ihre Entdeckung auf dem Gebiet der primären physiologischen und chemischen Sehvorgänge im Auge" ( gemeinsam mit Ragnar Granit und George Wald

31. Haldan Keffer Hartline - Prix Nobel De Physiologie Ou Médecine processus visuels physiologiques et chimiques primaires de l oeil http://www.nobelpreis.org/francais/medizin/hartline.htm

Suivant Rechercher Accueil Chimie ... Physique Haldan Keffer Hartline (Etats Unis) "Pour la découverte sur les processus visuels physiologiques et chimiques primaires de l'oeil" ( avec Ragnar Granit et George Wald

32. Entrez PubMed Nobel prize for George Wald, haldan keffer hartline and Ragner Granit in 1967for discoveries concerning the primary biochemical and physiological http://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=Retrieve&db=PubMed&list_uids=4

33. Entrez PubMed haldan keffer hartline 22 December 190318 March 1983. Granit R, Ratliff F.Publication Types Biography Historical Article MeSH Terms http://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=Retrieve&db=PubMed&list_uids=1

34. Hartline, Haldan Keffer --  Britannica Concise Encyclopedia - Your Gateway To hartline, haldan keffer body US physiologist. More results on hartline,haldan keffer when you join. 15 items. Subscribe to unlock this content today http://concise.britannica.com/ebc/article-9366668

Home Browse Store Help Search Britannica Concise Again Hartline, Haldan Keffer Concise Encyclopedia Article Page 1 of 1 Haldan Keffer Hartline born Dec. 22, 1903, Bloomsburg, Pa., U.S. died March 17, 1983, Fallston, Md. U.S. physiologist. He received his M.D. from Johns Hopkins University. Experimenting on horseshoe crabs, he was the first to record the electrical impulses sent by a single optic-nerve fibre. He found that when one of the eye's receptor cells is stimulated, others nearby are depressed, enhancing contrast and sharpening perception of shapes. He showed how simple retinal mechanisms constitute vital steps in the integration of visual information. In 1967 he shared a Nobel Prize with George Wald and Ragnar Arthur Granit var mm = [["Jan.","January"],["Feb.","February"],["Mar.","March"],["Apr.","April"],["May","May"],["June","June"],["July","July"],["Aug.","August"],["Sept.","September"],["Oct.","October"],["Nov.","November"],["Dec.","December"]]; To cite this page: MLA style: "Hartline, Haldan Keffer." Britannica Concise Encyclopedia http://concise.britannica.com/ebc/article-9366668

35. Wald, George --  Britannica Concise Encyclopedia - Your Gateway To All Britann Biographies of Ragnar Granit, haldan keffer hartline, and George Wald. Includes textof the presentation speech outlining their achievements in this field. http://concise.britannica.com/ebc/article-9382237

Home Browse Store Help Search Britannica Concise Again Wald, George  Concise Encyclopedia Article Page 1 of 1 George Wald born Nov. 18, 1906, New York, N.Y., U.S. died April 12, 1997, Cambridge, Mass. U.S. biochemist. vitamin A , the mechanisms of the photochemical reaction s in the rod cells that enable night vision, and the identification of the colour-sensitive pigments in the cone cells ( see photoreception retina ). He shared the Nobel Prize in 1967 with Haldan Keffer Hartline and Ragnar Arthur Granit . He was a prominent opponent of the Vietnam War. var mm = [["Jan.","January"],["Feb.","February"],["Mar.","March"],["Apr.","April"],["May","May"],["June","June"],["July","July"],["Aug.","August"],["Sept.","September"],["Oct.","October"],["Nov.","November"],["Dec.","December"]]; To cite this page: MLA style: "Wald, George." Britannica Concise Encyclopedia http://concise.britannica.com/ebc/article-9382237 APA style: Wald, George. ( Britannica Concise Encyclopedia . Retrieved http://concise.britannica.com/ebc/article-9382237 Britannica style: "Wald, George." Britannica Concise Encyclopedia http://concise.britannica.com/ebc/article-9382237

36. Haldan Keffer Hartline Biography .ms haldan keffer hartline. See also. Quotations. haldan keffer hartline (December22, 1903 March 17, 1983) was an American physiologist who was a cowinner http://haldan-keffer-hartline.biography.ms/

Haldan Keffer Hartline See also Quotations Haldan Keffer Hartline December 22 March 17 ) was an American physiologist who was a cowinner (with George Wald and Ragnar Granit ) of the Nobel Prize in Physiology or Medicine for his work in analyzing the neurophysiological mechanisms of vision. Hartline began his study of retinal electrophysiology as a National Research Council Fellow at Johns Hopkins University Baltimore , receiving his M.D. in . After attending the universities of Leipzig and Munich as an Eldridge Johnson traveling research scholar, he became professor of biophysics and chairman of the department at Johns Hopkins in . He joined the staff of Rockefeller University New York City , in as professor of neurophysiology. Hartline investigated the electrical responses of the retinas of certain arthropods, vertebrates, and mollusks because their visual systems are much simpler than those of humans and are thus easier to study. He concentrated his studies on the eye of the horseshoe crab (Limulus polyphemus). Using minute electrodes in his experiments, he obtained the first record of the electrical impulses sent by a single optic nerve fibre when the receptors connected to it are stimulated by light. He found that the receptor cells in the eye are interconnected in such a way that when one is stimulated, others nearby are depressed, thus enhancing the contrast in light patterns and sharpening the perception of shapes. Hartline thus built up a detailed understanding of the workings of individual photoreceptors and nerve fibres in the retina, and he showed how simple retinal mechanisms constitute vital steps in the integration of visual information.

37. Ragnar Granit - By Yrsa Neuman/Gunnar Damström At the same time British physiologist haldan keffer hartline conducted researchat the University of Philadelphia. In Pennsylvania Granit had the http://sfhs.eget.net/P_articles/Q_artiklar/Q_Granit.html

Ragnar Granit by Yrsa Neuman At the University of Philadelphia Ragnar Granit had large research facilities at his disposal and received generous grants for acquisition of research equipment. At the same time British physiologist Haldan Keffer Hartline conducted research at the University of Philadelphia. In Pennsylvania Granit had the opportunity to experimantally test his theory on the nervous layer of the retina and its importance for seeing. Ragnar Granit returned to Helsinki in 1934, and was shocked and saddened by the persecution of the Swedish language by the fennomans at the University of Helsinki. In his book he describes the hooligans hollering in the University aula and tarring Swedish language wall signs. Granit wished to research whether the retina had separate receptors for the different components of solar light, in other words about the way the eye sees color. Granit reached conclusions similar to those of the classics of color vision theory the early English physicist Thomas Young and the German physicist and physiologist Hermann von Helmholz : that the retina has three color sensitive layers. It turned out that the eye entails a kind of nerve center that prepares the information for the brain and for the actual way there. At the Carolinian Institute Granit focused his research on how the spinal marrow and the brain exert muscular control. He retired in 1967, the same year he shared the Nobel Price with Hartline and George Wald for his discoveries concerning the physiological and chemical foundations of vision. He passed away in 1991.

38. Biografia De Hartline, Haldan Keffer Translate this page hartline, haldan keffer. (1903-83) Científico norteamericano, n. en Bloomsburg (Pa.)y m. en Fallston. Especializado en biofísica, realizó estudios en el http://www.biografiasyvidas.com/biografia/h/hartline.htm

Inicio Buscador Las figuras clave de la historia Reportajes Los protagonistas de la actualidad Hartline, Haldan Keffer (1903-83) Científico norteamericano, n. en Bloomsburg (Pa.) y m. en Fallston. Especializado en biofísica, realizó estudios en el Lafayette College, la Universidad Johns Hopkins y en Leipzig y Munich, habiendo realizado actividades docentes en el Cornell University Medical College y en la Universidad de Pensilvania. En 1967 le fue otorgado el premio Nobel de Fisiología y Medicina junto a George Wald y Ragnar Granit. Los trabajos de investigación por los que Hartline había merecido tan alta distinción académica se referían a la transmisión de impulsos nerviosos del ojo al cerebro y el comportamiento de las fibras ópticas en las variaciones de la luz, para lo cual había tenido que emplear microelectrodos que aislaban las fibras y registraban sus reacciones al paso de la luz hasta la retina. Inicio Buscador Recomendar sitio

39. HARTline: Information From Answers.com haldan keffer hartline Brand alliances Ragnar Granit List of towns inWashington hartline, Washington Ybor City Paul Greengard 1967 in science http://www.answers.com/topic/hartline

showHide_TellMeAbout2('false'); Business Entertainment Games Health ... More... On this page: Wikipedia Mentioned In Or search: - The Web - Images - News - Blogs - Shopping HARTline Wikipedia HARTline HARTline logo HARTline provides public transportation for Hillsborough County Florida . The prefix of HARTline is an acronym that stands for "Hillsborough Area Regional Transit Authority". HARTline provides bus service and manages the TECO Line Streetcar System External links HARTline http://www.hartline.org/ TECO Line Streetcar System http://www.tecolinestreetcar.org/ This entry is from Wikipedia, the leading user-contributed encyclopedia. It may not have been reviewed by professional editors (see Mentioned In HARTline is mentioned in the following topics: Haldan Keffer Hartline Brand alliances Ragnar Granit List of towns in Washington ... George Wald Wikipedia information about HARTline This article is licensed under the GNU Free Documentation License . It uses material from the Wikipedia article "HARTline" More from Wikipedia Your Ad Here Jump to: Wikipedia Mentioned In Or search: - The Web - Images - News - Blogs - Shopping Send this page Print this page Link to this page Tell me about: Home About Tell a Friend Buzz ... Site Map

40. 1949: Information From Answers.com haldan keffer hartline b. Bloomsburg, Pennsylvania, December 22, 1903, d.Fallston, Maryland, March 17, 1983 resumes his study of reactions in a single http://www.answers.com/topic/1949

showHide_TellMeAbout2('false'); Arts Business Entertainment Games ... More... On this page: US Literature Wikipedia Mentioned In Or search: - The Web - Images - News - Blogs - Shopping In the year Archaeology A diver spots amphoras (Greek storage jars) on the sea bottom off the tiny island of Grand Conglou© a few kilometers outside the harbor at Marseilles, France. In 1952 the site is determined to be the wrecks of two ships from classical times. Mexican archaeologist Alberto Ruz Lhuillier [b. Paris, January 27, 1906, d. Montreal, Canada, August 25, 1979] excavates the previously untouched tomb of Pacal, eighth-century ce Maya ruler of Palenque. The tomb is filled with rich jade jewelry. The body wears a dramatic jade burial mask, reminding archaeologists of that of Tutankhamen. Astronomy Fred Lawrence Whipple [b. Red Oak, Iowa, November 5, 1906] suggests (correctly) that comets are "dirty snowballs," consisting of ice or ammonia ice and rock dust. See also 1577 Astronomy William Albert Hiltner [b. 1914, d. 1991] and John Scoville Hall [b. Old Lyme, Connecticut, June 20, 1908, d. October 15, 1991] discover polarization of light by interstellar matter. See also 1819 Astronomy Ralph Belknap Baldwin [b. 1912] formulates his meteor theory of the lunar surface. Before Baldwin, the most common explanation of the lunar craters had been volcanism. Baldwin demonstrates convincingly that the craters have been caused by impacts and not by volcanoes.

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